Levels of explanation in GWAS:

• https://www.cambridge.org/core/books/levels-of-analysis-in-psychopathology/entity-focus-applied-genetic-science-at-different-levels/C4182CB6CBFD6DA16E8BF25F64C19636
• https://www.journals.uchicago.edu/doi/pdf/10.1086/710519
• https://link.springer.com/article/10.1007%2Fs10519-018-9935-x - this paper explains implicit concepts underlying people's work on genetics well